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He is known for his portrayals of William Shakespeare in Shakespeare in Love (1998), for which he was nominated for the BAFTA Award for Best Actor in a Leading Role and the Screen Actors Guild Award for Outstanding Performance by a Male Actor in a Leading Role, Sir Robert Dudley in Elizabeth (1998), Commisar Danilov in Enemy at the Gates (2001), and Monsignor Timothy Howard in the second season of the TV series American Horror Story (2012–2013).He currently stars in the drama series The Handmaid's Tale (2017–present).His first professional stage appearance was in the West End in The Woman in Black, followed by A Month in the Country opposite Helen Mirren.Fiennes then became a member of the Royal Shakespeare Company for two seasons.His surname comes from the French village of Fiennes, Pas-de-Calais.
In 2002 he starred in the independent film, Killing Me Softly.
He then attended art school in Suffolk for a year, before joining the Young Vic Youth Theatre.
Fiennes subsequently trained for three years at London's Guildhall School of Music and Drama, graduating in 1993.
His nephew Hero Fiennes-Tiffin played Tom Riddle, young Lord Voldemort in Harry Potter and the Half-Blood Prince.
Fiennes was briefly educated in the Republic of Ireland, then at Swan School For Boys (now known as Leehurst Swan School, following a merger with another school), an independent school in Salisbury, before passing his 11 exam and continuing to Bishop Wordsworth's School, a voluntary-aided state day grammar school, in the Cathedral Close of the city.